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In angiosperms, the two male gametes that are formed by division of the generative cell. They migrate down the pollen tube behind the vegetative nucleus. When the pollen tube enters the embryo sac the tip of the tube breaks down to release the generative nuclei. On entering the nucellus and embryo sac they participate in the fertilization process. One fuses with the egg nucleus to form the zygote and the other usually fuses with the polar nuclei or definitive nucleus to form the primary endosperm nucleus.
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